Pcgf3 (polycomb group ring finger 3) - Chinchilla Research Resource Database
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Gene: Pcgf3 (polycomb group ring finger 3) Mus musculus
Analyze
Symbol: Pcgf3
Name: polycomb group ring finger 3
CRRD ID: 1551802
Description: Predicted to have metal ion binding activity. Involved in histone H2A-K119 monoubiquitination and inactivation of X chromosome by genetic imprinting. Localizes to PcG protein complex. Colocalizes with X chromosome. Is expressed in several structures, including branchial arch; central nervous system; limb; otocyst; and urinary system. Orthologous to human PCGF3 (polycomb group ring finger 3); PARTICIPATES IN histone modification pathway; INTERACTS WITH 1,2-dimethylhydrazine; 2,3,7,8-tetrachlorodibenzodioxine; bisphenol A.
Type: protein-coding
RefSeq Status: VALIDATED
Also known as: 2310035N15Rik; AI662857; D630042K08Rik; DON; DONG1; E430039C14; polycomb group RING finger protein 3; ring finger protein 3; RING finger protein 3A; RN; Rnf3; RNF3A
Orthologs:
Homo sapiens (human) : PCGF3 (polycomb group ring finger 3)  HGNC  Alliance
Rattus norvegicus (Norway rat) : Pcgf3 (polycomb group ring finger 3)  Alliance
Chinchilla lanigera (long-tailed chinchilla) : Pcgf3 (polycomb group ring finger 3)
Pan paniscus (bonobo/pygmy chimpanzee) : PCGF3 (polycomb group ring finger 3)
Canis lupus familiaris (dog) : PCGF3 (polycomb group ring finger 3)
Ictidomys tridecemlineatus (thirteen-lined ground squirrel) : Pcgf3 (polycomb group ring finger 3)
Sus scrofa (pig) : PCGF3 (polycomb group ring finger 3)
Chlorocebus sabaeus (African green monkey) : PCGF3 (polycomb group ring finger 3)
Heterocephalus glaber (naked mole-rat) : Pcgf3 (polycomb group ring finger 3)
more info ...
Latest Assembly: GRCm38 - Mouse Genome Assembly GRCm38
Position:
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm395108,609,082 - 108,654,842 (+)NCBI
GRCm385108,461,259 - 108,506,976 (+)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl5108,461,232 - 108,506,976 (+)EnsemblGRCm38mm10GRCm38
MGSCv375108,890,351 - 108,932,118 (+)NCBIGRCm37mm9NCBIm37
MGSCv365108,701,634 - 108,743,401 (+)NCBImm8
Celera5105,582,876 - 105,624,630 (+)NCBICelera
Cytogenetic Map5FNCBI
JBrowse: View Region in Genome Browser (JBrowse)
Model


Disease Annotations     Click to see Annotation Detail View

Gene-Chemical Interaction Annotations     Click to see Annotation Detail View
Gene Ontology Annotations     Click to see Annotation Detail View

Biological Process

Cellular Component

Molecular Function

Molecular Pathway Annotations     Click to see Annotation Detail View
Phenotype Annotations     Click to see Annotation Detail View

Mammalian Phenotype
References

Additional References at PubMed
PMID:10349636   PMID:10922068   PMID:11042159   PMID:11076861   PMID:12477932   PMID:12904583   PMID:14610273   PMID:15031102   PMID:15489334   PMID:16141072   PMID:16141073   PMID:17296600  
PMID:18287559   PMID:18799693   PMID:20059953   PMID:21267068   PMID:21677750   PMID:21873635   PMID:28049731   PMID:28596365   PMID:29054931   PMID:29220657   PMID:29727681   PMID:30186101  


Genomics

Comparative Map Data
Pcgf3
(Mus musculus - house mouse)
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm395108,609,082 - 108,654,842 (+)NCBI
GRCm385108,461,259 - 108,506,976 (+)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl5108,461,232 - 108,506,976 (+)EnsemblGRCm38mm10GRCm38
MGSCv375108,890,351 - 108,932,118 (+)NCBIGRCm37mm9NCBIm37
MGSCv365108,701,634 - 108,743,401 (+)NCBImm8
Celera5105,582,876 - 105,624,630 (+)NCBICelera
Cytogenetic Map5FNCBI
PCGF3
(Homo sapiens - human)
Human AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCh38.p13 Ensembl4705,748 - 770,640 (+)EnsemblGRCh38hg38GRCh38
GRCh384705,832 - 770,980 (+)NCBIGRCh38GRCh38hg38GRCh38
GRCh374699,537 - 764,428 (+)NCBIGRCh37GRCh37hg19GRCh37
Build 364689,573 - 754,428 (+)NCBINCBI36hg18NCBI36
Build 344689,572 - 754,255NCBI
Celera4654,586 - 701,073 (+)NCBI
Cytogenetic Map4p16.3NCBI
HuRef4674,466 - 735,220 (+)NCBIHuRef
CHM1_14699,310 - 761,729 (+)NCBICHM1_1
Pcgf3
(Rattus norvegicus - Norway rat)
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Rnor_6.0142,247,805 - 2,297,155 (-)NCBIRnor6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl142,247,808 - 2,297,155 (-)EnsemblRnor6.0rn6Rnor6.0
Rnor_5.0142,242,206 - 2,295,509 (-)NCBIRnor5.0Rnor_5.0rn5Rnor5.0
RGSC_v3.4141,824,819 - 1,839,263 (-)NCBIRGSC3.4rn4RGSC3.4
RGSC_v3.1141,784,275 - 1,835,105 (-)NCBI
Celera141,276,942 - 1,313,583 (-)NCBICelera
Cytogenetic Map14p22NCBI
Pcgf3
(Chinchilla lanigera - long-tailed chinchilla)
Chinchilla AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChiLan1.0 EnsemblNW_004955514307,332 - 350,039 (+)EnsemblChiLan1.0
ChiLan1.0NW_004955514307,331 - 350,039 (+)NCBIChiLan1.0ChiLan1.0
PCGF3
(Pan paniscus - bonobo/pygmy chimpanzee)
Bonobo AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
PanPan1.14746,147 - 805,443 (+)NCBIpanpan1.1PanPan1.1panPan2
PanPan1.1 Ensembl4746,137 - 805,443 (+)Ensemblpanpan1.1panPan2
Mhudiblu_PPA_v04802,990 - 874,063 (+)NCBIMhudiblu_PPA_v0panPan3
PCGF3
(Canis lupus familiaris - dog)
Dog AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
CanFam3.1 Ensembl391,676,496 - 91,708,754 (-)EnsemblCanFam3.1canFam3CanFam3.1
CanFam3.1391,673,424 - 91,708,751 (-)NCBICanFam3.1CanFam3.1canFam3CanFam3.1
Pcgf3
(Ictidomys tridecemlineatus - thirteen-lined ground squirrel)
Squirrel AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
SpeTri2.0NW_00493647722,349,166 - 22,373,256 (-)NCBISpeTri2.0SpeTri2.0SpeTri2.0
PCGF3
(Sus scrofa - pig)
Pig AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Sscrofa11.1 Ensembl8140,446 - 182,261 (+)EnsemblSscrofa11.1susScr11Sscrofa11.1
Sscrofa11.18140,435 - 189,796 (+)NCBISscrofa11.1Sscrofa11.1susScr11Sscrofa11.1
Sscrofa10.2880,086 - 128,768 (-)NCBISscrofa10.2Sscrofa10.2susScr3
PCGF3
(Chlorocebus sabaeus - African green monkey)
Vervet AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChlSab1.1 Ensembl2748,020,010 - 48,053,520 (-)Ensembl
ChlSab1.12748,015,976 - 48,077,688 (-)NCBI
Pcgf3
(Heterocephalus glaber - naked mole-rat)
Molerat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HetGla 1.0NW_00462475526,294,675 - 26,344,864 (-)NCBI

Position Markers
AI662857  
Mouse AssemblyChrPosition (strand)SourceJBrowse
GRCm385108,504,111 - 108,504,229UniSTSGRCm38
MGSCv375108,933,130 - 108,933,248UniSTSGRCm37
Celera5105,625,640 - 105,625,758UniSTS
Cytogenetic Map5FUniSTS
Whitehead/MRC_RH51331.38UniSTS


QTLs in Region (GRCm38)
The following QTLs overlap with this region.    Full Report CSV TAB Printer Gviewer
CRRD IDSymbolNameLODP ValueTraitSub TraitChrStartStopSpecies
11341716Rvfs3_mRift Valley fever susceptibility 3 (mouse)526236236138457140Mouse
1301524Lmb2_mlupus in MRL and B6 F2 cross (mouse)Not determined537424144112317856Mouse
4142064Tmc1m4_mTmc1 modifier 4 (mouse)Not determined554342835126924008Mouse
4142317Rgcs1_mretinal ganglion cell susceptible 1 (mouse)Not determined55230992108614761Mouse
26884395Humsd2_mhumerus midshaft diameter 2, 10 week (mouse)569900000132500000Mouse
27226719Tibw6_mtibia width 6, proximal, 16 week (mouse)572700000140000000Mouse
13506928Recrq8_mrecombination rate in male meiosis QTL 8 (mouse)574300000132500000Mouse
25314315Mlh1fc4_mMLH1 foci count 4 (mouse)574300000132500000Mouse
1357431Cia27_mcollagen induced arthritis 27 (mouse)Not determined575154687119880564Mouse
4141438Skmw13_mskeletal muscle weight 13 (mouse)Not determined576042404110042620Mouse
4141505Chlq14_mcirculating hormone level QTL 14 (mouse)Not determined576132927110133058Mouse
1301691Cdcs10_mcytokine deficiency colitis susceptibility 10 (mouse)Not determined576132927110133058Mouse
1300986Hdlq2_mHDL QTL 2 (mouse)Not determined576132927110133058Mouse
1300931Cocrb6_mcocaine related behavior 6 (mouse)Not determined576297205110297366Mouse
1301829Thyls2_mthymic lymphoma susceptibility 2 (mouse)Not determined576297205110297366Mouse
1301713Dbsty2_mdiabesity 2 (mouse)Not determined576297205110297366Mouse
13464249Ahl16_mage related hearing loss, early onset 16 (mouse)576297205110297366Mouse
10401246Bglu13_mblood glucose level 13 (mouse)Not determined579391781113391927Mouse
10755524Mono1_mmonocyte differential 1 (mouse)580701400114701400Mouse
1301419Cypr3_mcytokine production 3 (mouse)Not determined580871790114871944Mouse
10401245Bglu18_mblood glucose level 18 (mouse)Not determined582690609116690738Mouse
13464242Ahl17_mage related hearing loss, early onset 17 (mouse)582690609116690738Mouse
1300801Drb2_mdopamine receptor binding 2 (mouse)Not determined587239005121239199Mouse
1301399Cpfd3_mcerebellum pattern fissures (mouse)Not determined587239005121239199Mouse
1301126Bwem1_mbody weight day 30 males 1 (mouse)Not determined587239005121239199Mouse
10043958Chldq12_mcholesterol and HDL QTL 12 (mouse)Not determined587239005121239199Mouse
13464241Ahl18_mage related hearing loss, early onset 18 (mouse)587239005121239199Mouse
1357725Eae39_mexperimental allergic encephalomyelitis 39 (mouse)Not determined589271017118725963Mouse
1300968Skts4_mskin tumor susceptibility 4 (mouse)Not determined589271017124100176Mouse
1301362Prnr1_mprion resistance 1 (mouse)Not determined589271017146851125Mouse
11049562Lmr24f_mleishmaniasis resistance 24f (mouse)590173096124173175Mouse
1301457Cfsw2_mcystic fibrosis survival to weaning 2 (mouse)Not determined590413140124413285Mouse
1300993Hycdc_mhypercapnic duty cycle (mouse)Not determined590413140124413285Mouse
9587780Afw16_mabdominal fat weight QTL 16 (mouse)Not determined592682300126682419Mouse
1301974Chab7_mcholesterol absorption 7 (mouse)Not determined595085348129085464Mouse
1357644Egrd1_mearly growth rate, direct effect 1 (mouse)Not determined595286714129286920Mouse
1558774Lith17_mlithogenic gene 17 (mouse)Not determined595317683129317856Mouse
10402486Dipa1_mdrug induced psychomotor activation 1 (mouse)Not determined595317683129317856Mouse
13464244Ahl19_mage related hearing loss, early onset 19 (mouse)595317683129317856Mouse
1300940Actd3_mactivity-distance traveled 3 (mouse)Not determined596172519130172667Mouse
1300659Pcd8ts1_mp-glycoprotein positive CD8 T cell subset 1 (mouse)Not determined596883692130883925Mouse
1301858Smdq1_msegregation of mitochondrial DNA QTL 1 (mouse)Not determined596883692130883925Mouse
10045978Tir10_mtrypanosome infection response 10 (mouse)Not determined597871790113883925Mouse
1301042Elmaz1_melevated maze behavior 1 (mouse)Not determined597871790116738565Mouse
1301877Bbaa3_mB.burgdorferi-associated arthritis 3 (mouse)Not determined599307426112317856Mouse
4141519Hmtb5_mhemostasis and thrombosis bleeding time 5 (mouse)Not determined99801563126805880Mouse
12904945Tammq2_mtibialis anterior muscle mass QTL 2 (mouse)5100527900134527900Mouse
12904959Smmq1_msoleus muscle mass QTL 1 (mouse)5100527900134527900Mouse
12859290Criq2_mCitrobacter rodentium infection QTL 2 (mouse)5101194514135194514Mouse
1301624Aevm2_mautoimmune extremity vasculitis in MRL mice 2 (mouse)Not determined5101725767135725963Mouse
14746986Manh58_mmandible shape 58 (mouse)5102308218136308218Mouse
11532736Sluc33b_msusceptibility to lung cancer 33b (mouse)5102806972136807131Mouse
25314310Syncl4_msynaptonemal complex length 4 (mouse)5102900000115400000Mouse
25314317Sccor3_msynaptonemal complex length to mean MLH1 count ratio 3 (mouse)5104200000114600000Mouse
11528550Sluc33_msusceptibility to lung cancer 33 (mouse)5104239005120573921Mouse
1357886Diht_mdopamine-induced hypothermia (mouse)Not determined5104350872114963481Mouse
10412202Bbaa26_mB.burgdorferi-associated arthritis 26 (mouse)Not determined5104435822138557193Mouse
10412239Alpq7_malcohol preference QTL 7 (mouse)Not determined5104567876138567876Mouse
1357901Mdmsc2_mmodifier of muscularity 2 (mouse)Not determined5107413140112286920Mouse
4142473Chlq11_mcirculating hormone level QTL 11 (mouse)Not determined5107829235141829348Mouse
10043890Trigq4_mtriglyceride QTL 4 (mouse)Not determined5107829235141829348Mouse
13464243Ahl20_mage related hearing loss, early onset 20 (mouse)5107829235141829348Mouse

miRNA Target Status

Predicted Target Of
Summary Value
Count of predictions:3160
Count of miRNA genes:945
Interacting mature miRNAs:1344
Transcripts:ENSMUST00000046975, ENSMUST00000112597, ENSMUST00000138264, ENSMUST00000143973, ENSMUST00000151897
Prediction methods:Miranda, Rnahybrid, Targetscan
Result types:miRGate_prediction

The detailed report is available here: Full Report CSV TAB Printer

miRNA Target Status data imported from miRGate (http://mirgate.bioinfo.cnio.es/).
For more information about miRGate, see PMID:25858286 or access the full paper here.


Expression


Sequence


Reference Sequences
RefSeq Acc Id: ENSMUST00000046975   ⟹   ENSMUSP00000041790
RefSeq Status:
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm38.p6 Ensembl5108,461,232 - 108,506,976 (+)Ensembl
RefSeq Acc Id: ENSMUST00000112597   ⟹   ENSMUSP00000108216
RefSeq Status:
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm38.p6 Ensembl5108,461,232 - 108,489,224 (+)Ensembl
RefSeq Acc Id: ENSMUST00000138264   ⟹   ENSMUSP00000142465
RefSeq Status:
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm38.p6 Ensembl5108,461,313 - 108,486,171 (+)Ensembl
RefSeq Acc Id: ENSMUST00000143973
RefSeq Status:
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm38.p6 Ensembl5108,473,887 - 108,478,510 (+)Ensembl
RefSeq Acc Id: ENSMUST00000151897
RefSeq Status:
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm38.p6 Ensembl5108,486,220 - 108,496,290 (+)Ensembl
RefSeq Acc Id: NM_001377000   ⟹   NP_001363929
RefSeq Status: VALIDATED
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395108,609,125 - 108,654,842 (+)NCBI
GRCm385108,461,259 - 108,506,976 (+)NCBI
RefSeq Acc Id: NM_001377001   ⟹   NP_001363930
RefSeq Status: VALIDATED
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395108,609,125 - 108,654,842 (+)NCBI
GRCm385108,461,259 - 108,506,976 (+)NCBI
RefSeq Acc Id: NM_172716   ⟹   NP_766304
RefSeq Status: VALIDATED
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395108,609,125 - 108,654,842 (+)NCBI
GRCm385108,461,259 - 108,506,976 (+)NCBI
MGSCv375108,890,351 - 108,932,118 (+)RGD
Celera5105,582,876 - 105,624,630 (+)RGD
cM Map5 ENTREZGENE
Sequence:
RefSeq Acc Id: XM_030254794   ⟹   XP_030110654
RefSeq Status:
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395108,609,317 - 108,652,720 (+)NCBI
GRCm385108,461,503 - 108,504,473 (+)NCBI
Sequence:
RefSeq Acc Id: XM_036165475   ⟹   XP_036021368
RefSeq Status:
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395108,609,082 - 108,648,310 (+)NCBI
RefSeq Acc Id: XM_036165476   ⟹   XP_036021369
RefSeq Status:
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395108,609,317 - 108,648,310 (+)NCBI
RefSeq Acc Id: XM_036165477   ⟹   XP_036021370
RefSeq Status:
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395108,609,082 - 108,645,769 (+)NCBI
Reference Sequences
RefSeq Acc Id: NP_766304   ⟸   NM_172716
- Peptide Label: isoform 1
- UniProtKB: Q8BTQ0 (UniProtKB/Swiss-Prot)
- Sequence:
RefSeq Acc Id: XP_030110654   ⟸   XM_030254794
- Peptide Label: isoform X2
RefSeq Acc Id: NP_001363930   ⟸   NM_001377001
- Peptide Label: isoform 1
RefSeq Acc Id: NP_001363929   ⟸   NM_001377000
- Peptide Label: isoform 2
RefSeq Acc Id: ENSMUSP00000041790   ⟸   ENSMUST00000046975
RefSeq Acc Id: ENSMUSP00000142465   ⟸   ENSMUST00000138264
RefSeq Acc Id: ENSMUSP00000108216   ⟸   ENSMUST00000112597
RefSeq Acc Id: XP_036021368   ⟸   XM_036165475
- Peptide Label: isoform X1
RefSeq Acc Id: XP_036021370   ⟸   XM_036165477
- Peptide Label: isoform X3
RefSeq Acc Id: XP_036021369   ⟸   XM_036165476
- Peptide Label: isoform X1

Promoters
RGD ID:6887470
Promoter ID:EPDNEW_M7186
Type:initiation region
Name:Pcgf3_1
Description:Mus musculus polycomb group ring finger 3 , mRNA.
SO ACC ID:SO:0000170
Source:EPDNEW (Eukaryotic Promoter Database, http://epd.vital-it.ch/)
Experiment Methods:Single-end sequencing.
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm385108,461,277 - 108,461,337EPDNEW
RGD ID:6838006
Promoter ID:MM_KWN:43286
Type:CpG-Island
SO ACC ID:SO:0000170
Source:MPROMDB
Tissues & Cell Lines:3T3L1_Day0,   3T3L1_Day1,   BoneMarrow_0Hour,   BoneMarrow_2Hour,   BoneMarrow_4Hour,   Brain,   ES_Cell,   Kidney,   Liver,   Lung,   MEF_B4,   MEF_B6
Transcripts:ENSMUST00000112597,   OTTMUST00000042247,   OTTMUST00000042249
Position:
Mouse AssemblyChrPosition (strand)Source
MGSCv365108,890,191 - 108,890,692 (+)MPROMDB
RGD ID:6838007
Promoter ID:MM_KWN:43288
Type:Non-CpG
SO ACC ID:SO:0000170
Source:MPROMDB
Tissues & Cell Lines:BoneMarrow_4Hour,   Liver
Transcripts:OTTMUST00000042251
Position:
Mouse AssemblyChrPosition (strand)Source
MGSCv365108,914,319 - 108,914,819 (+)MPROMDB

Additional Information

Database Acc Id Source(s)
AGR Gene MGI:1916837 AgrOrtholog
Ensembl Genes ENSMUSG00000033623 Ensembl, ENTREZGENE, UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
Ensembl Protein ENSMUSP00000041790 ENTREZGENE, UniProtKB/Swiss-Prot
  ENSMUSP00000108216 UniProtKB/Swiss-Prot
  ENSMUSP00000142465 UniProtKB/TrEMBL
Ensembl Transcript ENSMUST00000046975 ENTREZGENE, UniProtKB/Swiss-Prot
  ENSMUST00000112597 UniProtKB/Swiss-Prot
  ENSMUST00000138264 UniProtKB/TrEMBL
Gene3D-CATH 3.30.40.10 UniProtKB/Swiss-Prot
InterPro RAWUL UniProtKB/Swiss-Prot
  Znf_RING UniProtKB/Swiss-Prot
  Znf_RING/FYVE/PHD UniProtKB/Swiss-Prot
  Znf_RING_CS UniProtKB/Swiss-Prot
KEGG Report mmu:69587 UniProtKB/Swiss-Prot
MGD MGI:1916837 ENTREZGENE
NCBI Gene 69587 ENTREZGENE
Pfam RAWUL UniProtKB/Swiss-Prot
PhenoGen Pcgf3 PhenoGen
PROSITE ZF_RING_1 UniProtKB/Swiss-Prot
  ZF_RING_2 UniProtKB/Swiss-Prot
SMART RING UniProtKB/Swiss-Prot
UniGene Mm.261218 ENTREZGENE
  Mm.486543 ENTREZGENE
UniProt A0A0G2JDR0_MOUSE UniProtKB/TrEMBL
  PCGF3_MOUSE UniProtKB/Swiss-Prot, ENTREZGENE
UniProt Secondary Q8BZT0 UniProtKB/Swiss-Prot